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Black-headed Antbird (Percnostola rufifrons) are found in shrubby borders, edges, gaps, and second growth in the northern Amazon basin and Guiana Shield. Sexes are indistinguishable and female-like early in life, washed brown-gray throughout with buffy-tipped coverts, until the preformative molt (FPF). These are the only known Amazonian species that exhibit a limited presupplemental (FPS) also known as the auxiliary preformative molt (FPX). Thus, this species' molt pattern is strikingly similar to Northern Cardinal (Cardinalis cardinalis) where the FPS is limited, while the subsequent FPF and definitive prebasic molts (DPB) are complete. FPS includes body feathers, and variable numbers of less coverts and med coverts. The minimum first cycle formative (M-FCF) males are black with white-edging to the wing coverts. The profile photo is of an M-FCF female followed by a series of synchronous age photos of male wings. 

Percnostola rufifrons

FCJ - First Cycle Juvenile

First cycle juvenile (FCJ ) individuals characterized by a single generation of loosely textured and gray-brown plumage. Coverts edged buff. Tail feathers noticeably more tapered than FAS individuals.

FCX - First Cycle Auxiliary Formative

First cycle Auxiliary Formative (FCX) individuals have replaced and female-like body plumage with noticeable molt limits between juvenal greater coverts and replaced less and med coverts. Sometimes these molt limits extend to the greater coverts. Sexes are often indistinguishable at this age, however, occasionally, males will replace one to several coverts that are black and edged white. FCX plumage is quickly replaced during the preformative molt (FPF).

FPF - First PreFormative

First preformative (FPF) molts are complete whereby juvenal remiges and rectrices are symmetrically and sequentially replaced by maturated plumage. FPF is most obvious in males whereby gray-brown juvenal plumage is replaced by black feathers. It is often more difficult to separate definitive prebasic (DPB) from FPF females because juvenal and female plumage is similar in color. In such cases, females can be aged as a minimum first cycle formative (M-FCF) or as an unknown molting individual (UPU). 

M-FCF - Minimum First Cycle Formative

Minimum first cycle formative (M-FCF) individuals are characterized by a single generation of plumage. Because the preformative molt is complete, it is often impossible to differentiate formative and definitive basic plumages. The WRP system accounts for this conundrum with the central "M" adjunct code, indicating that this bird is at a minimum in first cycle formative plumage. 

DPB - Definitive PreBasic

Definitive prebasic molt (DPB) is characterized by non-juvenal plumage (either formative or definitive basic plumage) being replaced by definitive basic plumage. Because the preformative molt is complete, and results in a formative plumage indistinguishable from the definitive basic plumage, it is often impossible to seperate SPB from DPB birds. Thus, individuals replacing maturated plumage with definitive basic feathers can be classified as DPB.

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